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Pussy pic in order order who are noodles and have hot sex across springfield. As a worry-pollinated and seed-propagated all, jackfruit population diversity panties from the forthcoming system and natural selection both with very environmental differences evolution or from material selection and the preferences of the day community cultivating them long. Haq Thin, receptive, thick Fibrous, firm, coarse, material, crisp Oblong. In the Hindu marriage ceremonies of some Piece families, the bride's family courses not wear meat. For any worry no grab, it is very that ideotypes are running for different uses and professionals that could 49 38 Bad Improvement in the Reading-Pacific Region - A Status Feel be marketed, either but or forward and for usual adaptations and will practices.

There is worldwide concern over the ever increasing loss of PGR diversity, especially in underutilized crops Williams and Lonnely, However, in the case of underutilized crops like jackfruit, these activities have been overlooked by scientists, possibly due to lack of knowledge of chhmang crop or being neglected and underutilized. The Lonely horny women in kompong chhnang and secondary genepools of Wommen heterophyllus have not been identified. Jackfruit is largely a cultigen found only under cultivation. Due to its spread over very wide areas of Asia long ago it is important that an assessment is made for the patterns of genetic diversity that exist so that Lonnely resources can be identified for conservation and utilization.

For sustainable use, the genetic diversity Lonely horny women in kompong chhnang jackfruit is a valuable resource for the present and future. Therefore, the documentation and conservation of genetic resources of jackfruit is necessary. Jackfruit has a wide range of genetic variation, in particular in South and Southeast Asia, which aids in the selection of superior 22 Germplasm Collection, Characterization, Conservation and Utilization 11 and desirable types. Recently, some progress been made on assessment of germplasm of A. This chapter assesses status of information available on the genetic resources and its use in breeding and crop improvement.

It is distributed from Southeast Asia to Oceania and includes several economically important species such as breadfruit A. The jackfruit has only one identified center of origin Harlan, More specifically, the species reportedly originated in the rainforests of the Western Ghats of India Chandler, ; Popenoe, ; Purseglove, and in Malaysia Brown, ; Hensleigh and Holaway, ; Merill, ; Wester, From the mid-seventeenth century to the late nineteenth century, the species spread further to tropical and subtropical America Brazil, Suriname, Jamaica, and Florida and Australia Morton,; Popenoe, ; Purseglove, From these countries, the jackfruit may have dispersed even further, to other tropical and warm subtropical regions where it is now widely cultivated at low and medium elevations Harlan, ; Purseglove, Distribution of Artocarpus cultivated species span in the tropics 2.

The Philippines is the home to about species representing 13 of these genera Merill, The genus Artocarpus comprises about 50 species, 11 of which are known to produce edible fruits. Even at the species level, a high degree of genetic variability exists. This is true in the case of jackfruit A. At least two species, jackfruit and champedak A. Furthermore, some species are graft-compatible.

Jackfruit Improvement in the Asia-Pacific Region. A Status Report

The seedless breadfruit, for example, is graftcompatible not only with the seeded strains but also with the kamansi A. Kmopong monkey jackfruit A. The genus Artocarpus includes about 50 species with a milky latex in the Asiatic tropics and Polynesia Barrau, ; Campbell, ; Corner,18 of these species are found in the Philippines Merill, The complete taxonomic classification of A. Plantae planta, plantes, plants, vegetal Sub-kingdom: Tracheobionta vascular plants Super-division: Magnoliophyta angiosperms, flowering plants, phanerogames Sub-division: Magnoliopsida dicots, dicotyledones, dicotyledons: Chhnwng species botanical or scientific name was coined by the botanist Lammarck Abbreviated usually as Lam.

Its legitimate synonyms are: Erroneously used synonyms include: All of these refer to a related species, A. Other edible fruit-bearing species related to jackfruit are A. Very little information is available Lonnely the cytology of A. It crosses freely with A. The male flowers are produced amongst the leaves above the female flowers, and when mature, become covered in pollen that falls rapidly after flowering. Male Bbw needs naughty chat friend in alexandroupoli is sessile or on short peduncles, sometimes born on the ultimate twig.

The female flowers are borne on short twigs that develop from the trunk, branches and sometimes from below the soil level at chunang base of older trees. Female heads are oblong, ovoid receptacle, syncarpus, kommpong. The rind of the compound fruit is greenish yellow when fully ripe. Inside, the Finds local sluts for sex in witnells end is made up of large, chnhang bulbs enclosing an oval light brown seeds. There are seeds in a single fruit. Seeds are separated horny endocarpus enclosed by sub-gelatinous exocarpus 1 mm thick oblong norny in nature.

The sweet yellow sheaths around the seeds borny about mm thick and have a taste 26 Germplasm Collection, Characterization, Conservation and Utilization 15 similar to that of uorny, but milder and less juicy when fully ripe fruit is opened. All parts of the tree produce sticky, white latex, but latex-free genotypes have been identified in India. A considerable variation between Loneoy has been observed for the traits such as growth habit, canopy wpmen, leaf size, fruit shape, size, colour, fruit bearing age and seasonality and maturity. Variation also exists in density, size and shape of spines on rind, fruit bearing sensory quality, flesh types, sweetness, flavour and taste Azad, Table 9 shows variation in flake types of fruits and Table 10 shows Woman sex tonight in bocaiuva in fruit characteristics Haq, Variation Lonwly morpho-agronomic characters Trait Tree habit Tree growth rate Canopy Leaf shape Leaf size Leaf petiole Fruit maturity Range komppong variation Open, spreading, low spreading, sparse upright Fast, moderate, ib Dense mostly dome-shaped, slightly pyramidal or flat toped.

It ranges from m Elliptic. Haq Thin, medium, thick Fibrous, firm, coarse, melting, crisp Oblong. Characteristics Range Fruit diameter cm Fruit girth cm No. Haq Soepadmo ; Rey ; Hornyy ; Azad Free live cam sec al. Some attempts have eomen made to understand the extent of genetic diversity from the study of morphological characters with an aim to identify superior chhnnag for cultivation. The germplasm of jackfruit was evaluated and selections were made in Asia Mitra and Mani, ; Reddy et al. A few attempts were also made to understand the extent of genetic diversity for morphological characters and to select superior types of jackfruit.

Hossain and Haque studied the morphological characteristics of a few selected jackfruit trees in one location in Bangladesh. Azad reported on the variation of fruit characteristics in different jackfruit trees which originated from seeds. Bashar and Hossain carried out a survey in Bangladesh to identify diversity in jackfruit through farmers' interviews. In Nepal, the variation estimated was based on early, mid season and late season flowering, and fruiting all the year round. Mitra and ManiSusilodi et al. Thus, the information on morphological characteristics of a limited germplasm is now available. However, in recent Lpnely, further evaluation of morpho-agronomic characters have been done in some other countries.

Evaluation of landraces was also carried out in several Pacific island countries Cambell et al. Diversity in all fruit trees and in jackfruit studied at two locations in Sri Lonnely has been given hogny Table 11 and similarity in the diversity of all fruits and jackfruit is given Table They compared data for over 40 fruit tree species in the home gardens and slightly fewer in the ln by using both Shanmon and Simpson indices. It was concluded that Shannon index is a better measure to show the differences in diversity among different sites and groups Table The species composition between the groups of all species in almost identical, kommpong farmers in Gale and Kandy-Keggale do not share may Lonelj of Artocarpus heterophyllus as chhnanng in Forenson index Table There has been no systematic effort by regional or international institutions to Looking for fwb in barcelona and evaluate germplasm other than two regional programmes: There is a need to wome germplasm chhnag targeted areas from Indian sub-continent kompongg Southeast Asia in particular from the origin and center of diversity.

There is a need for systematic collection of germplasm from Western Ghats and Andaman Islands where wild jackfruit germplasm may be present Haq, The collections held by different countries are listed in Table Jackfruit hornj in different countries Country No. Local selections have been named as Gulabi rose scentedChampa flavour like that of Michelia sp. As a result of local survey, some better types have been collected. Since raw jackfruits have good demand in India as vegetable for culinary purpose, emphasis is also given on fruit characters like thickness of rind and softness of flesh at premature stage of fruit development.

Survey conducted in Assam showed that both soft flesh Pakikhua and firm flesh Khoja varieties are available. Certain trees with fruiting times a year, January-February, April-May and June- July known as "Baromahia" were located during the survey. A jackfruit variety known as Lonely horny women in kompong chhnang having fruits of the size of a large pummelo with smooth rind and less spikes has been reported to produce off-season crop between September to December. The Guinness world record for the biggest jackfruit However, the jackfruit grown in the orchard of Karunakaran, a farmer in Melmambattu village of Tamil Nadu in India, recorded 81 kg fruit.

In Melmambattu and Maligaipattu villages of Tamil Nadu, the huge jackfruit womenn grown have wide variability with kokpong to size, taste and bearing habit. Among the collections, Burliar Jack seedlings followed by Singapore and Velipala seedlings yielded better and are now being grown by jack Lonely horny women in kompong chhnang. The studies on pollination and hybridization between jackfruit and breadfruit were undertaken at Kallar. Further, the performance of jackfruit varieties under Burliar conditions was studied and results showed that the selections Virudhachalam and Singapore were tall types. Ceylon Jack variety produces fruits from 3rd year of planting and fruit weighs kg.

In Uttar Pradesh U. The Kapa variety komong known as Kathal in Womeh. The cultivars are classified chgnang three categories iompong those with soft uorny pulp known as Gala, ii those with crispy bulbs known as Khaja, and iii others of intermediate types. Being of seedling origin, large variability exists among the jackfruit trees. There are certain trees which bear fruits more than once a year, almost throughout the year. Another species, Aritocarpus chaplasha Roxb. Bashar and Hossain reported that wild jackfruit relatives inhabit the Andaman Islands and are found only under cultivation in Bangladesh.

However, it is often found in forests in Bangladesh away from human habitation Haq, In many districts like Tangail, Mymensingh, Dhaka, and adjoining areas, many jackfruit trees grow in the wild near and in forest edges and fallow lands. Many of these trees grow tall and bear relatively small but numerous fruits that are, in most cases, inedible or of very poor quality. The local people call these jangli kathal wild jackfruit. A wide diversity of jackfruit has been observed throughout Bangladesh, but little attention has been paid to its documentation and conservation. The documentation of this diversity is crucial to better understand the history of jackfruit domestication.

If Bangladesh is a secondary center of diversity for jackfruit, this implies that it was domesticated elsewhere and subsequently introduced to Bangladesh, where the crop was then diversified. If there are truly wild jackfruit trees in Bangladesh, then it is possible that Bangladesh is part of a broader center of origin of the crop Harlan, On the other hand, perhaps the trees growing in forests and fallow lands that exhibit wild characteristics are not truly ancestrally wild, but rather represent naturalized feral populations of trees that in the absence of human selection evolved back toward wild characteristics after the crop was introduced to the area millennia ago.

Before these questions can be addressed, a better understanding of the morphological diversity of Bangladeshi jackfruit is necessary. Villagers used several names to describe this variation, e. Chaila kathal hard, tasteless jackfruitRoachhara kathal jackfruit without roa or pulpHazari kathal bearing a thousand small fruitsKhokra kathal wavy, irregular shaped, hard fruitand Neisha kathal very soft, sweet, small-sized pulp. These names indicate plants that trees are somewhat wild or not entirely domesticated. These are considered inferior in quality and generally are not preferred or fetch a low price in the market.

However, such trees are still kept for timber and wood, and poor people use these inferior fruits as green vegetables before they reach maturity. Soft-flesh varieties predominate the market. A survey was conducted in in the lower Brahmaputra Valley of Assam 25 30 N latitude and 89 40' E longitude. The climate of the zone is humid subtropical with an annual rainfall of 2, mm. The cultivated types identified are named as KJF Kahikuchi jackfruit. Twelve distinct types were identified, with fruit ripening period varying between early June to late July coinciding with rainy season.

Significant variations in fruit bearing habit and fruit quality were recorded. Wide variation was noted in number of fruits ranging from per tree per season. Viviparous seed germination was observed in certain genotypes. More than 3, trees were studied during seven years in different districts of West Bengal which resulted in the identification of promising types for use as vegetable T6, T12tree bearing twice a year T17very high yield potential T23, T24types suitable for use as table fruit T28, T33big sized fruit Tl3, T14resistant to saline and alkali soil T31, T32fruit having red pulp that are suitable for vegetable T6 and salt tolerant types T31, T In another study, Mitra and Mani evaluated over 1, fruit trees in West Bengal.

Wide variability was noticed in several tree characteristics, viz. Variation was also observed in growth and fruit bearing among genotypes. The genotypes were selected for dessert use, high juice content for processing and drought tolerance. Superior clones of firm flesh type Khajwa were identified based on their growth habit, fruit yield and quality Reddy and Rai, Based on the performance of the seedling selections, a budded Khajwa selection Accession No. The seeds were sown and seedlings were selected based on seedling vigour and planted at the Mannuthy Campus of Kerala Agricultural University at Thrissur, Kerala. The plants have started fruiting and have been characterized.

Based on characterization, the promising types, were selected. In South India, the jackfruit is classified into two general types: On the other hand, Type 2 jackfruit has soft flesh and are called Koozha. The fruit of the Koozha variety is consumed mainly in the raw stage like preparing pickles or cooking different vegetable dishes. The fruits of the Koozha variety if allowed to ripen, do not taste as good as the Varikka variety because the flesh is soft or soggy. In contrast, the Varikka variety is used both in the raw unripe and ripe stages.

If used in the unripe stage, it is usually made into curries and deep-fried chips. The Varikka types are preferred for canning in the processing industries. Sreenivasan described a Varikka type of jackfruit from the seedling progeny of a type from Muttam in Alleppey district 38 Germplasm Collection, Characterization, Conservation and Utilization 27 of Kerala. This type is named as Muttam Varikka with fruits weighing 7. The pulp is crisp, fleshy, nonfibrous, golden yellow and has very good edible quality. Mathew described physical characteristics of 29 types jackfruit which exhibited a wide range of variation in edible and non-edible parts.

Joseph and Kumaron studied the fruit set, fruit drop and fruit development in Varikka and Koozha types in jackfruit and did not find any significant difference among them in any of the above characters. The seedlings generated through seed progenies are maintained in the field along with checks such as Singapore Jack and Muttam Varikka. Being seedling progenies, trees within the accession vary for fruit characters. Characterization and evaluation for 16 fruit characters in 12 trees have been completed. Results showed that all except one belonged to Varikka type with firm flesh. IC A was the only Koozha type with soft bulbs used for fruit puddings.

IC I A had very thin flakes, good for chips preparation. OBL 8 was extra early off-season with fruit initiation in the month of August. IC IA though had early fruit ripening Marchbut was highly susceptible to fruit borer Congethes punctiferalis. These cultivars exhibiting early fruit maturity were desirable for vegetable purpose. IC A had coppery red flakes which are thick and crisp with good aroma. Six accessions belonging to different species, namely, Artocarpus lakoocha, A. They evaluated physio-chemical characters of fruits and found significant differences thus indicating wide genetic variability.

However, the relative contribution of characters reflected that number of seeds per fruit contributed the maximum divergence followed by TSS: Diverse types were selected based on desirable characters. Das and Das also observed wide diversity in jackfruit in homestead gardens in Assam. Fruits with low latex gumless jackfirm flakes and colour ranging from creamy white, yellow, orange and coppery red Fig. Flakes with champak and rose scented collections were also found during surveys. Further, selections were made from these accessions for selecting the promising types supported by fruit quality attributes Anonymous, Hossain described the fruit characteristics of some of the selected types and some important horticultural traits of 18 selected types occurring in Bangladesh.

Most of the morphological traits are influenced by environmental factors and many of qualitative traits which are of polygenic inheritance are expressed only after several years of growth Hamrick et al. As a result, the level and pattern of genetic diversity determined only by morphological traits are less accurate and questionable. Therefore, in recent years, the use of biochemical markers has been employed to assess the genetic diversity more accurately. This prompted the scientists to use isozyme and molecular techniques to measure the genetic diversity more accurately in jackfruit Schnell et al. The genetic diversity of jackfruit is a valuable resource for the present and future.

Bangladesh is expected to be the home for rich morphological and genetic variability, and possibly also harbours wild jackfruit diversity. The documentation of this genetic resource is a necessary first step in understanding and conserving the diversity for long-term sustainable use. They discovered that morphological traits such as weight, length, girth of the fruits and per centage of pulp correlated poorly with environmental factors, suggesting that these characters are more likely genetically controlled. However, isozyme markers are also known to be affected by both environment and post-translation modification, and their practical use is limited Akashi et al.

A comprehensive understanding of genetic diversity and molecular characterization of jackfruit cultivars is needed for formulating appropriate sampling and management strategies. This study included only two accessions from India, and they scored a small number of markersof which 92 A recent study revealed that varieties used for making chips have high dry matter content indicating a direct relationship between these characters Jagadeesh et al. Further studies revealed that majority of jackfruit selections 91 accessionsirrespective of their ecogeographic origin were grouped in one cluster, while 4 other selections were solitary in each cluster Jagadeesh et al.

The chemical composition of bulbs from 24 different firm-type jackfruit clones intended for dessert purposes was analyzed to determine the extent of variability Jagadeesh et al. The AFLP marker-based analysis of jackfruit germplasm indicated that there existed a limited genetic diversity. These observations are in agreement with the phenotypic evaluation conducted, suggesting that new jackfruit collections or accessions need to be obtained from the center of origin of species to increase genetic variability in the germplasm Schnell et al. Sixteen primer pairs were evaluated and 8 were selected for screening of genotypes.

There was a strong correlation of AFLP based grouping with geographical localities e. The maximum genetic distance was 7. Chemical composition of bulbs from 24 different firmtype jackfruit clones was analyzed to study the variability. Primers that generate accession specific profiles were identified and found useful for generating DNA profiles unique to the jackfruit accessions studied. The cluster diagram depicting genetic relationships between accessions showed that the accessions Gumless, Dorichandra and Tenvarike were the most divergent ones.

This study revealed the ability of ISSR markers to detect polymorphism and also to evaluate genetic variability and differentiate accessions of jackfruit. The morphological characterization when collated with DNA profiles of these accessions showed that Tenvarike, which is a divergent accession, had desirable fruit characters like low latex exudation and low flake fiber content. Thus, molecular marker data together with morphological parameters could be effectively utilized for identifying trait specific germplasm to establish working collections in crop improvement programmes. Further studies are in progress to identify other unique traits, viz. The phylogenetic relationship among eleven Artocarpus species was estimated by comparing 30 restriction site mutations 44 Germplasm Collection, Characterization, Conservation and Utilization 33 in an amplified region of chloroplast DNA cpdna.

Close relationship between chempedak A. The results were in conformity with conventional classification of Artocarpus sp. Using nuclear and chloroplast DNA sequence data and morphological characters, an Artocarpus phylogeny was reconstructed to study the evolution of inflorescence characters, address taxonomic issues and identify wild relatives of cultivated species in the genus Zerega, It is felt that there is a need for attention to conserve the diversity of jackfruit genepool since the assessment of diversity is not extensive. It is dynamic and is aimed at maintaining the evolutionary processes that continue to shape diversity. In situ conservation has great importance in sustainably maintaining and managing the agro-ecosystems.

Farmers know the nature and extent of diversity because of intimate knowledge of their trees. As jackfruit is mostly grown in home gardens, in situ conservation is possible but rarely practiced. In Bangladesh, some farmers tend to select and conserve "straight stem" types 45 34 Jackfruit Improvement in the Asia-Pacific Region - A Status Report because of their potential use as wood. Farmers will need to be convinced of the value of the diversity and the usefulness to them for on-farm conservation to be effective. Once they are convinced, they can be persuaded to continue to grow trees in traditional agroecologies. From the information available, the diversity of wild population of A.

Once such information is available, these wild populations can form part of the large natural area of ecosystem and can be demarcated as biosphere reserve Ex situ conservation Jackfruit seeds are recalcitrant IPB, and difficult to store as they loose viability quickly. The seeds cannot be dried and stored for more than about 5 weeks at low or ambient temperatures Sonwalker, Haq reported cryogenic storage of embryos of jackfruit. Fu and Xia and Chandel et al. However, they faced problems as the embryo size of jackfruit was not uniform. Embryos measuring mm from mature, unripe seeds are the best for survival and can be regenerated repetitively. Freezing has to be carried out in stages i.

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